The first section is that of the Bovinae, which includes buffaloes, bison and oxen.
Unlike the Bovinae, there are frequently glands in the feet; and the upper molar teeth differ from those of that group in their narrower crowns, which lack a distinct inner column.
The characteristics of the sub-family Bovinae, or typical section of the family Bovidae, are given in the article Bovidae; for the systematic position of that family see Pecora.
From all other Bovinae they differ by the triangular section of their horns.