Primitively there are four perradial tentacles, to which may be added four interradial, or they may become very numerous and are then scattered evenly round the margin, never arranged in tufts or clusters.
The transparent prolonged into the four long tissues allow the enteric cavities and oral arms, perradial in position.
The sixteen tentacles of the scyphistoma disappear, and in the place of the four perradial and four interradial tentacles, the eight tentaculocysts of the adult are formed as outgrowths of the subumbral margin, independently of the tentacles of the scyphistoma (Friedemann).
The ephyra has a flat, disk-shaped body, with eight marginal lobes (four perradial, four interradial); a tentaculocyst is lodged in a deep notch at the apex of each lobe.
With sixteen marginal lobes, four rhopalia and twelve tentacles; the rhopalia are perradial in position, corresponding to the angles of the stomach.