Satyrus (431-387), the successor of Spartocus, established his rule over the whole district, adding Nymphaeum to his dominions and laying siege to Theodosia, which was a serious commercial rival by reason of its ice-free port and direct proximity to the cornfields of the eastern Crimea.
A motley host, made up out of the tribes bordering on the Black Sea and the Caspian, hovered round his small army, but failed to hinder him from laying siege to the town.
But Memnon died at the critical moment whilst laying siege to Mytilene and the great plan collapsed.